Mona Lisa's smile: The mystery has finally been solved after 500 years

Leonardo da Vinci’s famous portrait of Lisa Gherardini, titled Mona Lisa, is one of the most-discussed works of art ever, partly thanks to the subject’s ambiguous look.

According to a recent study by the University of Freiburg, though, the answer to whether she is “sad” or “happy” is simple: her expression is unequivocally “happy”.

Twelve participants were shown nine black and white photos of the Mona Lisa, eight of which had been digitally manipulated at the mouth; four made the model look happier, the other four sadder.

After shuffling the photos and showing them to each participant 30 times, the team found that

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